• Humans, housecats.
  • Other than humans, surplus killing has been observed among zooplankton, damselfly naiads, predaceous mites, martens, weasels, honey badgers, wolves, orcas, red foxes, leopards, lions, spotted hyenas, spiders, brown and black and polar bears, coyotes, lynx, mink, raccoons, dogs, and house cats.

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